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Will a turbo 400 transmission fit on 1978 trans am 403?

Yes, a Turbo 400 transmission is a common and appropriate choice for a 1978 Trans Am with a 403 cubic inch engine. The 403 was frequently paired with a Turbo 400 in that era. However, you'll need to ensure you have the correct bellhousing to mate the transmission to the engine. The bellhousing is specific to the engine and transmission combination.